Maize (Zea mays) Intercropping with Legumes Enhances Growth, Dry Matter and its Forage Yield under Deficit Irrigation

 

Muhammad Aslam1*, Muhammad Naeem1, Abdul Rehman2,3, Muhammad Mubashar Zafar4, Rashid Iqbal1, Muhammad Ayaz Shahzad1, Rana Muhammad Ikram Khan1 and Javed Iqbal1

1Department of Agronomy, Faculty of Agriculture and Environment, The Islamia University of Bahawalpur, 63100, Pakistan

2Zhengzhou Research Base, State Key Laboratory of Cotton Biology, Zhengzhou University, Zhengzhou 450000, China

3Institute of Cotton Research, Chinese Academy of Agricultural Sciences, Anyang 455000, China

4State Key Laboratory of Cotton Biology, Key Laboratory of Biological and Genetic Breeding of Cotton, The Ministry of Agriculture, Institute of Cotton Research, Chinese Academy of Agricultural Science, Anyang, 455000, Henan, China

*For correspondence: aslam1302@gmail.com; aslam1302@yahoo.com

Contributed equally to this work and are co-first authors

Received 25 July 2020; Accepted 04 September 2020; Published 10 December 2020

 

Abstract

 

Climate change is one of the serious threats to agriculture and livestock. Climate change induced and reduced water shortage lowers production of food and fodder crops. In order to investigate the effects of deficit irrigation on forage yield of maize (Zea mays L.), cowpea (Vigna unguiculata L.), and guar (Cyamopsis tetragonoloba L.) grown as sole or intercropping of maize with cowpea and guar, the fields experiments were conducted during spring 2016 and 2017 in randomized complete block design (RCBD) with split plot arrangement. These crops were further exposed to water deficit by skipping irrigation at crop stages following maize growth scale at 15 days (V4), 30 days (V6), 45 days (V9), and 60 days (V12) after sowing along with normal irrigation. Maize intercropped with guar had maximum leaf area, plant height land equivalent ratio and monetary advantage index at normal irrigation but did not differ significantly under deficit irrigation. Thus, maize intercropped with guar was found more productive and beneficial with respect to sole under deficit irrigation. © 2021 Friends Science Publishers

 

Key words: Cowpea; Forage; Guar; Intercropping; Maize; Yield

 


Introduction

 

Sustainable agriculture based on the provision of food security, enhanced quantity and quality of agriculture produces to satisfy the demand of increasing population (Eskandari 2012). Livestock though the mainstay of agriculture sector has remained a neglected sector for years. Various factors are responsible for low productivity of dairy animals regarding milk and meat production. The non-provision of quality forages is the single main factor that squelches the animal’s productivity (Iqbal et al. 1999). Moreover, the availability of quality fodder to the livestock is contributed to sustainable milk and meat production (Ginwal et al. 2019). Therefore, in order to keep the livestock’s productivity up, it is much needed to increase supply of quality forage/fodder (Iqbal and Iqbal 2015).

Forages are considered most reliable and nutritious source of animal feed resource. The term forage represents all the plant materials in succulent and green form while the term fodder represents the dried form of feed such as silage and hay. Cereals forages such as millet (Panicum miliaceum L.), maize (Zea mays L.), and sorghum (Sorghum bicolor L.), are important source of animals feed but have less protein contents. Compared to the cereals, legume forages such as soybean (Glycine max L.), cow pea (Vigna unguniculata L.), cluster bean (Cyamopsis tetragonoloba L.) are more protenious but their yield per unit area is a matter of concern (Bhagmal et al. 2009). Growing of legumes with cereals crops is productive and evolves as profitable cropping system over solitary cropping (Evans et al. 2001). Growing of mixed crop would enhance the production on the given area, and economic resources utilization (Marer et al. 2007). Legumes fix the atmospheric nitrogen, improve soil fertility, and are also a good source of nitrogen for cereal crops (Manna et al. 2003). Growing of cereals with legumes is best way for the availability of nitrogen to the neighbor crop (Connolly et al. 2001). When legumes are intercropped with cereals, it improves the utilization of available resource for maximizing yield (Khonde et al. 2018).

Water is a limiting factor in dry areas. Deficit irrigation has been evolved a way not only to save the water but an important tool to get optimum yield. The effect of deficit irrigation is non-significant if applied on less sensitive growth stages of the crop (Moutonnet 2002). When maize crop receives three irrigations with the depth of 150 mm for each at vegetative, tasseling and grain filling stages, it would produce higher yield (Fentawa 2006). Maize can be intercropped with legumes to feed the animals at any stage without risk of any ingredient i.e., prussic acid and oxalic acid (Dahmardeh et al. 2009).

When cereals like wheat and barley (Hordeum vulgare L.) are intercropped with legumes like fababean (Vicia faba L.), and rapeseed (Brassica napus L.) under deficit irrigation, the yield of the cereals is not affected (Amanullah et al. 2020). When maize is intercropped with legumes under deficit irrigation, the yield of maize is also not significantly affected (Sani et al. 2014).

Various studies have been conducted to explore yield/economic advantages of the intercropping but to find a suitable combination of cereal and legumes to get optimum yield and forage quality under deficit irrigation has not been explored to its full potential. Therefore, to explore this area of research, three spring fodders viz., maize, cowpea, and guar were grown as sole crops and intercropping of maize with cow pea and guar under deficit irrigation and the impact of deficit irrigation was evaluated on the forage yield of the crops.

 

Materials and Methods

 

Experiment was carried out at experimental area Department of Agronomy, Faculty of Agriculture and Environment, The Islamia University of Bahawalpur, Pakistan (Latitude, 29° 23ʹ 60.00” N, Longitude, 71° 40ʹ 59.99” E). The soil of experimental area was analyzed before sowing the crops. The soil was found to be sandy loam type. The soil samples were taken from 0-15 cm and 16-30 cm depth. The soil physico-chemical properties of experimental area were analyzed at Soil and Water Testing Laboratory Regional Agriculture Research Institute Bahawalpur, Pakistan during 2016 and 2017 are given in Table 1.

The seeds of three spring fodders i.e. maize variety ‘Neelam’, local variety of guar ‘desi’, and local variety of cowpea ‘rawaan’ were collected from Ayub Agricultural Research Institute, Faisalabad, Pakistan and sown on February 15, 2016 and 2017, as sole maize, cowpea, and guar and intercropping of maize with cowpea and guar respectively, maintaining seed ratio of 70:30 for each combination (Azim et al. 2000).

The sole crops and their combinations were subjected to four water deficit irrigation regimes along with control applied at different interval days in the following manners: Normal Irrigation, V4= Four leaves development stage (15 days after sowing), V6= Six leaves development stage (30 days after sowing), V9= Nine leaves development stage (45 days after sowing), and V12=Twelve leaves development stage (60 days after sowing). The treatments were replicated three times using randomized complete block design (RCBD) under split plot design with net plot size of 3 m × 6 m. A fine seed bed was prepared by four ploughing followed by planking. Seeds were sown through hand drill on ridges maintaining row to row distance of 30 cm and legumes intercropped in alternate rows of maize. Fertilizer was applied at 112.50 kg N and 50 kg P2O5 ha-1 in each plot. The ½ of the nitrogen and whole phosphorous was applied as basal dose and remaining nitrogen was applied at succeeding irrigation. Seed rate for maize, cowpea and guar was used at the rate of 100, 30, and 50 kg ha-1 respectively. Maize crop was harvested at heading stage approximately 75 days after sowing (at least 35% dry matter) for green fodder. Legume fodders were harvested at early pod formation stage (8 weeks after sowing) (Azim et al. 2000; Iqbal et al. 2006).

Data regarding average environmental temperature and rainfall during field experiment in 2016 and 2017 was recorded respectively (Table 2).

 

Growth and yield parameters

 

Leaf area (cm) per plant was measured with the help of formula; LA= LW × A. Where LA is leaf area, L is leaf maximum length, and W is leaf maximum width and A is constant respectively. The value of the constant (A) is 0.75 (Montgomery 1911). A quadrant with dimension 1 m × 1 m was used while taking the sample from each plot. Number of plants (stand density; m-2) was calculated in area (Khan et al. 2014). Height of harvested plants (cm) of each crop was measured at maturity by taking five plants from each plot selected randomly with the help of meter rod (Khan et al. 2014). The randomly selected plants in the given area were separated and counted number of leaves per plant (Khan et al. 2014).

 

Competition indices

 

Land equivalent ration was measured with following formula: LER: (LER maize + LER legume). Where LER maize: (Yml /Ym) and LER legume: Ylm / Yl. Where Ym was yield of sole maize crop and Yl was yields of sole legumes crops, respectively. Yml was yield of maize intercrop and Ylm was yield of legumes intercrops respectively (Machet et al. 1997). The value of relative crowding coefficient (K) was calculated with following formula: K: (Kmaize × Klegume), Where Kmaize: Yml × Zlm/ (Ym - Yml) × Zml), and Klegume: Ylm × Zml / (Yl - Ylm) × Zlm). Where Zml and Zlm are the proportions of maize and legume in the mixture respectively (Dhima et al. 2007). Aggressivity (A) was measured as: Amaize

Table 1: Physico-chemical analysis of experimental soil

 

Description

2016

2017

Values

Status

Values

Status

Depth

0-15 cm

16-30 cm

0-15 cm

16-30 cm

Texture

sandy loam

Sandy loam

Sand percentage

41

38

39

37

Silt percentage

35

36

36

37

Clay percentage

24

26

25

26

Chemical analysis

0-15 cm

16-30 cm

0-15 cm

16-30 cm

EC

1.30 dS m-1

1.28 dS m-1

Normal

1.34 dS m-1

1.29 dS m-1

Normal

pH

8.7

8.9

Alkaline

8.6

8.4

Alkaline

Organic matter

0.93%

0.91%

Deficient

0.98%

0.95%

Deficient

P

16.90 mg kg-1

16.71 mg kg-1

Deficient

17.12 mg kg-1

17.09 mg kg-1

Deficient

N

0.016 mg kg-1

0.017 mg kg-1

Deficient

0.017 mg kg-1

0.018 mg kg-1

Deficient

K

125 mg kg-1

126 mg kg-1

Sufficient

126 mg kg-1

127 mg kg-1

Medium

EC: Electric conductivity; P: Phosphorous; N: Nitrogen; K: Potassium

 

Table 2: Average monthly temperature and rainfall during experimental duration (2016 and 2017)

 

Years

Months

Temperature (0C)

Rainfall (mm)

2016

February

20

12.2

March

25

14.1

April

30

16.5

May

32

18.1

2017

February

12

4.2

March

20

12.3

April

25

10.3

May

16

8.5

(Source: Arid Zone Research Institute, Bahawalpur, Pakistan)

 

= (Yml / Ym × Zml) – (Ylm / Yl × Zlm) and Alegume = (Ylm / Yl × Zlm) – (Yml / Ym × Zml) (Dhima et al. 2007). Competitive ratio (CR) was calculated with formula: CRmaize = (LERmaize / LERlegume) (Zlegume-maize / Zmaize-legume), and CRlegume = (LERlegume / LERmaize) (Zmaize-legume / Zlegume-maize) (Tsubo et al. 2005). Actual yield loss index (AYLI) was calculated as: AYL = AYLmaize + AYLlegume. Where AYL maize={(Yml /Zml) (Ym/ Zm)}−1 and AYL legume={(Ylm/ Zlm) (Yl /Zl)}−1 (Banik and Sharma 2009). Monetary advantage index (MAI) describes economic advantages of intercropping. It can be calculated by using following formula: MAI: (value of combined intercrops) (LER-1) / LER. Higher value of MAI results in profitable intercropping (Ghosh 2004). Intercropping advantage (IA) describes economic feasibility of intercropping and measured as: IAmazie: AYLmazie × Pmaize where IAlegume: AYLlegume × Plegume. Commercial values of legumes and maize are denoted by Pmaize and Plegume respectively (Banik et al. 2000). After harvesting plants from every plot with the help of sickle, their (t ha-1) weight was measured by spring balance (Iqbal et al. 2006). The dry matter yield was calculated with the help of formula; Dry matter (%): Dry weight/Fresh weight x 100 and Dry matter yield: Forage yield of related crop × dry matter (%) of that crop (Iqbal et al. 2006).

 

Statistical analysis

 

Data were analyzed statistically by using Fisher’s analysis of variance techniques using STATISTIX software and the differences among the treatment means were compared according to Least Significant Difference (LSD) at 5% probability (Steel et al. 1997).

 

Results

 

Growth and yield parameters

 

Leaf area per plant, stand density, green forage and dry matter yield of sole maize and intercrops were affected significantly by different levels of irrigation during both years respectively. Maximum leaf area per plant was exhibited by maize intercropped with guar at normal irrigation during both years while minimum leaf area per plant was exhibited by sole maize for irrigation skipped 45 and 60 days after sowing, respectively. Sole maize achieved maximum stand density, green forage and dry matter yield at normal irrigation with respects to its intercrop (Table 3).

Leaf area per plant, plant height, stand density, green forage and dry matter yield of cowpea grown as sole and intercropped with maize were affected significantly at each irrigation level during both years respectively. Maximum leaf area per plant was achieved by cowpea intercropped with maize at normal irrigation while sole cowpea resulted in minimum leaf area per plant at same level of irrigation. Maximum plant height was exhibited by sole cowpea at normal irrigation and

Table 3: Effect of different irrigation regimes on growth and yield parameters of maize grown as sole and intercropped with cowpea and guar

 

Leaf area (cm2) per plant

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

585.30d

551.23de

604.35cd

517.18e

598.24d

606.40e

623.25de

633.45c-e

626.54de

612.35e

Maize + cowpea

676.33ab

685.41ab

704.88ab

704.00ab

664.78bc

754.52ab

705.65ab

706.80ab

696.45bc

707.25ab

Maize + guar

729.81a

723.41ab

720.24ab

704.23ab

684.33ab

773.22a

733.68ab

689.12b-d

708.20ab

707.88ab

LSD value at 5%

64.16

56.76

Plant height (cm)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

215.89a

208.70ab

210.00ab

199.90ab

219.56a

249.80a

217.41b

205.37bc

216.38b

215.22b

Maize + cowpea

196.41ab

205.05ab

212.53ab

193.43ab

197.49ab

213.63b

215.78b

210.38b

209.50bc

211.23b

Maize + guar

199.20ab

207.00ab

183.44ab

205.48ab

208.31ab

208.04bc

208.73bc

181.98c

208.20ab

213.74b

LSD value at 5%

31.93

27.27

Number of leaves per plant

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

15a

13ab

12ab

12ab

13ab

16a

14ab

13ab

12b

14ab

Maize + cowpea

14ab

13ab

11b

12ab

14ab

15ab

13ab

14ab

13ab

13ab

Maize + guar

13ab

12ab

12ab

13ab

13ab

12b

13ab

13ab

15ab

15ab

LSD value at 5%

3.53

3.45

Stand density ( plants m-2)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

26a

21b-e

22a-c

23ab

22a-c

30a

24ab

23b

22b-d

23b

Maize + cowpea

18b-f

17c-f

16d-f

15ef

16ef

18c-e

16d-e

15e

16e

15e

Maize + guar

15f

15f

14f

16ef

19b-f

15e

14e

16e

15e

15e

LSD value at 5%

4.85

4.87

Green forage yield ( t ha-1)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

40.63a

34.257bc

33.83bc

34.45b

33.95bc

40.39a

31.63bc

33.027b

31.29bc

30.78b-d

Maize + cowpea

32.28bc

29.84d-e

31.16bc

28.73e

28.60d-e

29.90b-e

26.67d-f

28.49c-e

23.75f

26.06ef

Maize + guar

32.79b-d

30.80b-e

30.50c-e

32.24b-e

28.76e

29.38b-e

28.94b-e

29.61b-e

27.35d-f

27.47c-f

LSD value at 5%

3.85

3.78

Dry matter yield ( t ha -1)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

13.15a

10.20b

8.98bc

8.83bd

8.27c-e

12.16a

8.92bc

7.93cd

8.25bc

9.45b

Maize + cowpea

7.24d-e

7.38c-e

7.01e

8.04c-e

6.82e

6.36e

5.03f

5.89ef

5.72ef

6.55e

Maize + guar

8.01c-e

7.190de

6.87e

6.80e

6.60e

6.71de

6.70de

6.87de

6.80de

6.47e

LSD value at 5%

1.79

1.18

Where I1, I2, I3, and I4 indicate numbers of irrigation skipped after 15, 30, 45, and 60 days after sowing keeping I0 normal irrigation with no skipping. Values sharing same letters differ non-significantly (P > 0.05)

 

minimum plant height was achieved by cowpea intercrop for irrigation skipped 15 and 30 days after sowing, respectively. Maximum stand density, green forage and dry matter yield were exhibited by sole cowpea at normal irrigation while minimum for irrigation skipped 15 and 30 days after sowing while minimum green forage yield for irrigation skipped 30 and 60 days after sowing and dry matter yield for irrigation skipped 45 days after sowing by cowpea intercropped with maize respectively (Table 4).

Leaf area per plant, plant height, stand density, green forage and dry matter yield of sole guar and intercrop were affected significantly by all irrigation levels during both years, respectively. Maximum leaf area per plant and plant height were achieved by guar intercrop and sole at normal irrigation while minimum leaf area per plant for irrigation skipped 15 and 30 days after sowing and minimum plant height for irrigation skipped 60 and 45 days after sowing was achieved by sole guar respectively. Sole guar had stand density, green forage and dry matter yield at normal irrigation. The lowest stand density for irrigation skipped 45 and 30 days after sowing and lowest green forage and dry matter yield for irrigation skipped 60 days after sowing were achieved by guar intercrop. Maximum number of leaves per plant was achieved by sole guar at normal irrigation and minimum number of leaves per plant was exhibited by guar intercrop for irrigation skipped 15 and 45 days after sowing respectively (Table 5).

 

Competition indices

Table 4: Effect of different irrigation regimes on growth and yield parameters of cowpea grown as sole and intercropped with maize

 

Leaf area (cm2) per plant

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Cowpea

597.1d

554.95d

685.57a-c

624.67b-d

611.39cd

623.00d

642.39cd

717.48ab

676.30b-d

618.90d

Cowpea + maize

719.54a

722.67a

707.47ab

704.19ab

685.49a-c

755.44a

710.74a-c

707.47 a-c

695.49 a-c

707.46 a-c

LSD value at 5%

57.30

79.99

Plant height (cm)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Cowpea

162.00a

137.20c

137.02c

141.24bc

159.76ab

162.18a

137.20a-c

135.29a-c

153.01a-c

158.14ab

Cowpea + maize

135.58c

130.39c

146.30a-c

128.68d

133.39cd

128.42c

135.75a-c

132.48bc

135.76 a-c

140.43 a-c

LSD value at 5%

16.04

26.88

Number of leaves per plant

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Cowpea

80a

57bc

60bc

64bc

70ab

77a

61bc

73ab

66a-c

75ab

Cowpea + maize

68ab

65b

61bc

67ab

52c

69ab

67a-c

64 a-c

71ab

53c

LSD value at 5%

13.60

17.85

Stand density (plants m-2 )

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Cowpea

65a

59a

57a

60a

61a

60a

52a

54a

53a

51a

Cowpea + maize

41b

30b

38b

38b

40b

36b

32b

31b

34b

35b

LSD value at 5%

10.54

10.01

Green forage yield (t ha -1)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Cowpea

15.24a

12.58ab

12.96a

12.10a

13.54ab

16.14a

14.02ab

15.51ab

13.42b

14.47ab

Cowpea + maize

6.20bc

5.80c

5.30c

6.01bc

5.90c

7.10bc

6.80c

6.40c

5.98c

6.10c

LSD value at 5%

2.54

2.38

Dry matter yield (t ha -1)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Cowpea

4.01a

3.17a-c

3.13a-c

3.05a-c

3.50ab

5.12a

3.60b-d

3.91bc

3.58b-d

3.85bc

Cowpea + maize

2.68bc

2.37cd

2.17c

2.01d

2.05d

2.85c-e

2.19e

2.13e

2.10e

2.30c

LSD value at 5%

0.83

1.05

Where I1, I2, I3, and I4 indicate numbers of irrigation skipped after 15, 30, 45, and 60 days after sowing keeping I0 normal irrigation with no skipping. Values sharing same letters differ non-significantly (P > 0.05)

 

Maize showed dominance in maize: cowpea and maize: guar intercropping combinations as it showed higher values of land equivalent ratio (LER), relative crowding co-efficient (K), aggressivity (A), competition ratio (CR), actual yield loss index (AYLI), intercropping advantage (IA) at each irrigation level than cowpea and guar intercrops during both years respectively. The highest values of LER were recorded for maize-guar intercropping with guar at normal irrigation and minimum land equivalent ratio (LER) were recorded for irrigation skipped 30 and 15 days after sowing for maize-cowpea intercropping, respectively. It showed the economic advantages of maize-guar intercropping over maize-cowpea intercropping. Moreover, LER values of maize-guar intercropping did not differ significantly to each other at each irrigation level. Higher values of relative crowding co-efficient (K) for maize: guar intercropping were observed at normal irrigation while maize-cowpea resulted in minimum K values for irrigation skipped 15 days after sowing respectively that showed better performance over maize-cowpea intercropping. Different values of aggressivity (A) indicate that companion crops did not show equal competition and maize showed higher value to dominate over other intercrops. Higher values of A of maize-cowpea intercropping were recorded at normal irrigation as compared to A values of maize-guar intercropping that achieved minimum for irrigation skipped 15 and 45 days after sowing, respectively. Maximum values of competition ratio (CR) of maize were recorded at normal irrigation that showed its dominance over intercrops. Moreover maize-cowpea intercropping achieved higher values of CR at normal irrigation while minimum CR values of maize-guar intercropping were observed for irrigation skipped 30 and 60 days after sowing, respectively. Higher positive values of actual yield loss index (AYLI) of maize were observed at each irrigation level as compared to negative AYLI values of intercrops while maize-guar intercropping showed maximum values of AYLI at normal irrigation as compared to the minimum AYLI values of maize-cowpea intercropping for irrigation skipped 15 and 30 days after sowing respectively. Overall all positive values of AYLI indicate the economic advantages of intercropping. Likewise, maize resulted in to higher positive values of intercropping advantage (IA) at all irrigation levels as compared to negative IA values of intercrops and maize-guar intercropping resulted in to higher values of IA at normal irrigation as compared to the minimum IA values of maize-cowpea for irrigation skipped 30 and 45 days after sowing respectively. Over all positive values of IA showed the beneficial effects of intercropping. Maize-guar intercropping

Table 5: Effect of different irrigation regimes on growth and yield parameters of guar grown as sole and intercropped with maize

 

Leaf area (cm2) per plant

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Guar

703.20ab

563.19cd

616.43c

569.99cd

621.73c

735.67a

670.38cd

607.79e

607.80e

708.34a-c

Guar + maize

719.54a

690.67ab

707.47ab

704.19ab

685.49a-c

711.01ab

680.87bc

628.00e

633.22de

710.15 a-c

LSD value at 5%

48.04

32.75

Plant height (cm)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Guar

178.46a

164.29bc

168.88a-c

160.07bc

156.73c

184.67a

172.67ab

166.43ab

164.33b

173.33ab

Guar + maize

174.37a-c

166.50bc

170.96bc

172.34bc

169.80bc

185.21a

177.33ab

175.33ab

178.67ab

183.00ab

LSD value at 5%

20.31

20.47

Number of leaves per plant

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Guar

319.33a

313.33ab

304.67ab

292.67ab

288.00ab

317.00a

315.33ab

310.67ab

297.67ab

294.67ab

Guar + maize

286.00ab

263.00b

284.0ab

289.33ab

279.67ab

285.67ab

280.67ab

284.00ab

278.33ab

283.67ab

LSD value at 5%

55.99

44.56

Stand density (plants m-2)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Guar

38a

29b

28bc

28bc

26bc

45a

35b

32bc

33bc

34b

Guar + maize

21bc

18c

19bc

17c

19bc

24cd

21cd

20d

22cd

23cd

LSD value at 5%

9.89

7.10

Green forage yield (t ha -1)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Guar

21.48a

16.79b

16.92b

15.80b

15.05b

22.62a

19.86b

17.64c

17.80bc

16.35c

Guar + maize

9.35c

7.90c

7.80c

7.63c

7.05c

9.15de

9.00de

8.90e

8.16e

7.92e

LSD value at 5%

2.02

1.92

Dry matter yield (t ha -1)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

 

 

 

 

 

 

 

 

 

 

Guar

5.50a

5.10a

5.01b

4.84ab

4.53a-c

6.65a

5.57a

5.87a

5.86a

5.36a

Guar + maize

3.74b-d

3.65b-d

2.55b-d

3.50c-d

3.16d

3.01b

2.97b

2.99b

2.84b

2.73b

LSD value at 5%

1.33

1.54

Where I1, I2, I3, and I4 indicate numbers of irrigation skipped after 15, 30, 45, and 60 days after sowing keeping I0 normal irrigation with no skipping. Values sharing same letters differ non-significantly (P > 0.05)

 

 

resulted in the highest values of monetary advantage index (MAI) at normal irrigation while minimum MAI values of maize-cowpea intercropping were achieved for irrigation skipping 15 days after sowing respectively that resulted in higher economic advantages with respect to maize-cowpea intercropping. Moreover, the values of MAI did not significantly affect at each irrigation level and resulted in the economic advantages of intercropping under deficit irrigation (Table 6).

 

Discussion

 

Leaf area per plant, stand density, green forage and dry matter yield of sole maize and intercrops were significantly affected during both years (Table 3).

 

Table 6: Effect of different irrigation regimes on competition indices of maize intercropped with cowpea and guar

 

Land equivalent ratio (LER)

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

0.81c

0.80c

0.82c

0.81c

0.82c

0.81c

0.80c

0.77c

0.78c

0.79c

Cowpea

0.48d

0.50d

0.46de

0.39e

0.51d

0.59d

0.48e

0.53e

0.53e

0.52e

Guar

0.64d

0.58de

0.54de

0.58de

0.50e

0.58c

0.57c

0.54c

0.59c

0.54c

Maiz + Cowpea

1.29b

1.30b

1.28b

1.20b

1.33ab

1.40a

1.28b

1.30b

1.31b

1.31b

Maize + Guar

1.45a

1.38ab

1.36ab

1.39ab

1.32ab

1.39a

1.37a

1.31ab

1.37a

1.34ab

LSD at 5%

0.06

0.05

Relative crowding co-efficient (K)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

0.90a

0.92a

0.91a

0.86a

0.94a

0.93a

0.86a

0.88a

0.86a

0.90a

Cowpea

0.19c-e

0.13de

0.22c

0.20c-e

0.21cd

0.15f-h

0.13g

0.23b

0.18e

0.17e

Guar

0.57b

0.36bc

0.34bc

0.38bc

0.51bc

0.30b

0.29b

0.25b

0.30b

0.31b

Maiz + Cowpea

0.17c-e

0.12e

0.20cd

0.17c-e

0.19c-e

0.14g

0.11h

0.21d

0.16e-h

0.15f-h

Maize + Guar

0.51bc

0.33bc

0.31bc

0.33bc

0.47bc

0.28b

0.25b

0.22c

0.26b

0.28b

LSD at 5%

0.07

0.06

Aggressivity (A)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

0.57a

0.56a

0.57a

0.57a

0.57a

0.57a

0.57a

0.56a

0.56a

0.57a

Cowpea

0.21c

0.15d-e

0.14e

0.14e

0.16d

0.14e

0.14e

0.13e

0.16d

0.16d

Guar

0.18c

0.21c

0.22c

0.19c

0.18c

0.17c

0.17c

0.18cd

0.18cd

0.18cd

Maiz + Cowpea

0.45b

0.41b

0.43b

0.43b

0.42b

0.45d

0.42b

0.42b

0.41b

0.40b

Maize + Guar

0.39b

0.37b

0.35b

0.37b

0.39b

0.40b

0.40b

0.39b

0.38b

0.39b

LSD at 5%

0.04

0.05

Competition ratio (CR)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

0.82ab

0.80b

0.81ab

0.81ab

0.82a

0.82a

0.81ab

0.81ab

0.80ab

0.81ab

Cowpea

0.39i

0.50f

0.46h

0.48gh

0.51f

0.48hi

0.46hi

0.42i

0.50h

0.51g

Guar

0.60e

0.69b-d

0.71a-c

0.64d

0.67c

0.63d

0.57ef

0.58ef

0.55f

0.59ef

Maiz + Cowpea

0.70a-c

0.68cd

0.76bc

0.74bc

0.69cd

0.59g

0.74b-e

0.72c-e

0.69c

0.68c

Maize + Guar

0.61e

0.51f

0.49f

0.53d

0.57f

0.65c-e

0.62d

0.60e

0.61d

0.59ef

LSD at 5%

0.11

0.14

Actual yield loss index (AYLI)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

0.82ab

0.88ab

0.92a

0.81ab

0.96a

0.93a

0.73bc

0.87ab

0.88ab

0.94a

Cowpea

-0.43f

-0.52f

-0.37e

-0.42f

-0.37e

-0.38k

-0.42l

-0.50e

-0.37k

-0.24j

Guar

-0.12d

-0.26e

-0.15d

-0.20d

-0.24d

-0.26j

-0.29j

-0.28d

-0.27j-l

-0.32k

Maiz + Cowpea

0.50de

0.35d

0.55cd

0.39de

0.59cd

0.55d

0.31h

0.28i

0.51e

0.70b

Maize + Guar

0.77bc

0.61c

0.73bc

0.61c

0.72bc

0.67c

0.44g-i

0.48f

0.61c

0.62c-e

LSD at 5%

0.10

0.12

Intercropping advantage (IA)

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maize

1167.30a

910.50bc

1090.40ab

1103.80ab

1159.20a

1197.80a

1020.30ab

1032.50ab

1094.70ab

1144.30a

Cowpea

-539.40g

-459.20g

-651.70h

-515.70g

-467.70g

-470.10f

-463.50f

-249.30f

-561.00h

-499.50g

Guar

-144.80f

-192.60f

-324.20fh

-249.90f

-307.20g-i

-334.70e

-334.10e

-407.60ef

-350.20e

-370.70e

Maiz + Cowpea

627.90d

451.30e

438.70e

558.10de

691.40cd

727.70cd

556.80de

783.20c

533.70d

644.80de

Maize + Guar

1022.50ab

717.90bc

766.20c

853.90b

852.00b

863.10b

686.20c

624.90e

744.50c

773.60c

LSD at 5%

195.87

158.86

Monetary advantage index (MAI)

 

 

2016

2017

I

I

I

I

I

I

I

I

I

I

Maiz + Cowpea

10860d

12410cd

15516ab

12691b-d

11971cd

12925b-d

9548d

11869cd

11900cd

12329cd

Maize + Guar

16307a

15581a

14402a-c

14569a-c

14412a-c

15589a

12728c

15152a-c

15911a-c

16640ab

LSD at 5%

3305.6

4145.6

Where I1, I2, I3, and I4 indicate numbers of irrigation skipped after 15, 30, 45, and 60 days after sowing keeping I0 normal irrigation with no skipping. Values sharing same letters differ non-significantly (P > 0.05)

 

 

The higher values of leaf area per plant were recorded for maize-guar intercropping at normal irrigation. The higher leaf area is due to the beneficial effects of legumes intercrop over the companion crops as the former help to fix atmospheric nitrogen, increases over all productivity of the intercrops and higher leaf area contributed to maximum interception of light, utilized in photosynthesis which resulted in maximum forage yield (Nasrollahzadeh and Koohi 2014; Ginwal et al. 2019). The maximum stand density, green forage and dry matter yield were achieved by the sole maize during the same years at same irrigation level that might be due to the higher seed rate used for sole maize as compared to low seed rate with 70:30 for maize and cowpea intercrop respectively. This resulted in higher plant population, green forage and dry matter yield for sole maize as compared to maize intercrop (Khan et al. 2012). The leaf area per plant and plant height obtained under normal irrigation were not significantly differed as compared to under deficit irrigation. Sole cowpea and guar achieved maximum values of leaf area per plant, stand density, green forage and dry matter yield during both years as compared to intercrops. The higher leaf area of cowpea and guar was achieved due to non-competing effect of the intercrops (Abate and Alemayehu 2018; Yang et al. 2018). While higher stand density, green forage and dry matter yield was due to higher seed rate used for sole cowpea and guar as compared to the seed rate with 70:30 in intercropping with maize as higher seed rate contributed to maximum plant population and biological yield for sole cowpea and guar as compared to intercrops (Khan et al. 2012) (Table 3 and 4).

Land equivalent ratio (LER) measure the effectiveness of intercropping over mono-cropping to utilize environmental resources and its value greater than one, equal to one or less than one indicate more yield, same or less yield respectively (Khonde et al. 2018). The maize-guar intercropping resulted in maximum LER during both years and showed its advantages over maize-cowpea intercropping that might be due to effective and efficient utilization of natural resources like land and light (Yang et al. 2018). The relative dominance of one species over the other in a mixture is known as relative crowding co-efficient (K) and crop with higher value of K, the more dominant effect it would have over companion crop (Takim 2012). The results revealed that the partial K values of maize were greater than the partial K values of cowpea and guar during both years which showed competitive advantage of the maize over intercrops and maize intercropped with guar resulted in maximum K values as compared to its intercropping with cowpea (Khonde et al. 2018). The relative competitive ability of one crop as compared to other when cultivated in combination with that crop can be described with the help of aggressivity (A). If the value of A is either zero (0), or positive (above zero), and/or negative (less than zero) then both crops would be equally competitive, or one crop is dominant or weak in competition respectively (Jan et al. 2016). During both years of the experiment, the partial values of A of maize were greater than intercrops that showed the dominance of maize over cowpea and guar (Imran et al. 2011). It is clearly shown from the experiment that partial values of CR of maize were greater than intercrops during both years which resulted in competitive advantage of maize over cowpea and guar showing higher ability to utilize environmental resources efficiently (Javanmard et al. 2014). The proportionate yield advantage (gain) or disadvantage (loss) of intercrops as compared to sole crops is described as actual yield loss index (AYLI). It is derived from the experiment maize showed partial positive values of AYLI that resulted in yield advantages while partial values of AYLI of intercrops were negative that showed the yield disadvantages. Maize-guar resulted in maximum values of AYLI during both years of the experiment as compared to maize-cowpea intercropping (Takim 2012). The economic feasibility of intercropping is termed as intercropping advantage (IA). It is observed from the experiment that maize showed positive values of IA as compared to intercrops which showed negative values. However, over all positive IA values were recorded which showed the advantages of intercropping over solitary cultivation. Maize intercropped with guar resulted in maximum values of IA during experimental duration that resulted in to more feasible option of intercropping as compared to guar (Takim 2012). Monetary advantage index (MAI) describes economic advantages of intercropping. Maize-guar intercropping resulted in maximum values of MAI as compared to MAI values of maize-cowpea intercropping that showed its higher economic advantage over maize-cowpea intercropping under deficit irrigation during both years (Tofa et al. 2017) (Table 6).

 

Conclusion

 

This research study concludes that maize intercropped with cowpea and guar reduced the yield as compared to their sole cultivation but overall productivity in terms of leaf area, plant height, land equivalent ratio, and monetary advantage index were recorded in maize: guar intercropping under deficit irrigation and it did not differ significantly with respect to normal irrigation. So, maize: guar intercropping was found a suitable cropping combination under hot climate where farmer face scarcity of irrigation water.

 

Author Contributions

 

MA and MN planned, conduct, and write the initial results/draft of the experiment, AR and MMZ reviewed the results, RI, MAS, RMIK and JI statistically analyzed the data and made illustrations.

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